Woolly mammoths (Mammuthus primigenius) were among the most abundant cold adapted species during the Pleistocene. Their once large populations went extinct in two waves, an end-Pleistocene extinction of continental populations followed by the mid-Holocene extinction of relict populations on St. Paul Island, a small island in the middle of the Bering Sea, 5,600 years ago, and on Wrangel Island, a remote Arctic refuge off the coast of Siberia, 4,000...
